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Hollier, J.; Stöckli, E.; Wesener, T.; Mesibov, R.; Decker, P.; Sierwald, P. (2020). An annotated list of the millipede (Diplopoda) species described by Johann Carl. Revue suisse de Zoologie. 127(1): 183-240.
381271
10.35929/RSZ.0015 [view]
Hollier, J.; Stöckli, E.; Wesener, T.; Mesibov, R.; Decker, P.; Sierwald, P.
2020
An annotated list of the millipede (Diplopoda) species described by Johann Carl
Revue suisse de Zoologie
127(1): 183-240
Publication
Johann Carl described 390 species or subspecies of millipedes during his long career at the Muséum d’histoire naturelle de Genève, studying specimens collected on his own expeditions and on those of other, mainly Swiss, naturalists. The type specimens deposited in Geneva are enumerated, and the type specimens that could be located in other collections in other museums are reported. Lectotypes are designated for 17 species: Brachyspirobolus aequatorialis (Carl, 1909), Carlopeltis alatus (Carl, 1914), Mastigorhacus alatus (Carl, 1912), Salpidobolus annulipes (Carl, 1912), Erythrhacus arietis (Carl, 1912), Argentocricus bernardinensis (Carl, 1918), Trigoniulus bitaeniatus Carl, 1912, Desmocricus conjunctus Carl, 1918, Polylepis elberti Carl, 1912, Trigoniulus incommodus Carl, 1912, Stenobolus insularis Carl, 1918, Salpidobolus lateralis (Carl, 1912), Dinematocricus lombokensis (Carl, 1912), Prepodesmus longipes (Carl, 1913), Arostrophus mertoni (Carl, 1912), Salpidobolus moenensis (Carl, 1912) and Eucarlia velox (Carl, 1912). Polydesmus carli nom. nov. is introduced as a replacement name for Polydesmus japonicus Carl, 1902, a junior homonym of Polydesmus japonicus Peters, 1864. Odontopyge johanncarli nom. nov. is introduced as a replacement name for Odontopyge fasciata Carl, 1905, a junior homonym of Odontopyge fasciata Attems, 1896.
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Agathodesmus baccatus (Carl, 1926) (additional source)
Aphelidesmus ambiguus Carl, 1914 (additional source)
Argentocricus bernardinensis (Carl, 1918) (additional source)
Batodesmus acceptus Carl, 1914 (additional source)
Brachyspirobolus aequatorialis (Carl, 1909) (additional source)
Calymmodesmus andinus Carl, 1914 (additional source)
Canacophilus acutangulus Carl, 1926 (additional source)
Carlopeltis alatus (Carl, 1914) (additional source)
Chondrodesmus attemsi Carl, 1914 (additional source)
Cryptocorypha areata (Carl, 1932) (additional source)
Cyrtodesmus ammonites (Carl, 1914) (additional source)
Desmoxytes bicolor (Carl, 1902) (additional source)
Doratogonus annulipes Carl, 1917 (additional source)
Erythrhacus arietis (Carl, 1912) (additional source)
Harpurostreptus matarae Carl, 1941 (additional source)
Harpurostreptus montivagus Carl, 1941 (additional source)
Hyleoglomeris alticola (Carl, 1912) (additional source)
Ktenostreptus acceptus Carl, 1941 (additional source)
Leptodesmus augustus Carl, 1914 (additional source)
Leptostreptus caudatus Carl, 1941 accepted as Leptostreptus caudiculatus (Karsch, 1881) (additional source)
Lophostreptus bicolor Carl, 1909 (additional source)
Lophostreptus ptilostreptoides Carl, 1909 (additional source)
Mastigorhacus alatus (Carl, 1912) (additional source)
Microspirobolus aequatorialis Carl, 1909 accepted as Bukobolus aequatorialis (Carl, 1909) accepted as Brachyspirobolus aequatorialis (Carl, 1909) (additional source)
Neocordyloporus asperus Carl, 1905 (additional source)
Nesorthomorpha armata (Carl, 1902) (additional source)
Odontopyge johanncarli Hollier, Stöckli, Wesener, Mesibov, Decker & Sierwald, 2020 (status source)
Opisthoporodesmus bacillifer Carl, 1912 (additional source)
Picrogonopus gracillimus Carl, 1941 (additional source)
Polydesmus americanus Carl, 1902 accepted as Pseudopolydesmus pinetorum (Bollman, 1888) (taxonomy source)
Polydesmus carli Hollier, Stöckli, Wesener, Mesibov, Decker & Sierwald, 2020 (status source)
Polydrepanum asperrimum (Carl, 1932) (additional source)
Poratophilus distinctus Carl, 1917 accepted as Zinophora distincta (Carl, 1917) (additional source)
Poratophilus junodi Carl, 1917 accepted as Zinophora junodi (Carl, 1917) (additional source)
Poratophilus similis Carl, 1917 accepted as Zinophora similis (Carl, 1917) (additional source)
Prepodesmus ornatus (Peters, 1864) (additional source)
Pseudopolydesmus pinetorum (Bollman, 1888) (additional source)
Salpidobolus annulipes (Carl, 1912) (additional source)
Sholaphilus albidus Carl, 1932 (additional source)
Spirobolellus ambiguus Carl, 1926 (additional source)
Spirostreptus angustifrons Carl, 1905 (additional source)
Spirostreptus frater Carl, 1909 accepted as Gonoplectus frater (Carl, 1909) (additional source)
Spirostreptus inconstans Carl, 1914 accepted as Hyloecostreptus inconstans (Carl, 1914) (additional source)
Spirostreptus malayus Carl, 1909 accepted as Gonoplectus malayus (Carl, 1909) (additional source)
Spirostreptus minor Carl, 1909 accepted as Thyropygus minor (Carl, 1909) accepted as Remulopygus minor (Carl, 1909) (additional source)
Spirostreptus multiannulatus Carl, 1909 (additional source)
Spirostreptus neglectus Carl, 1909 accepted as Remulopygus neglectus (Carl, 1909) (additional source)
Spirostreptus pulcherrimus Carl, 1909 accepted as Ktenostreptus pulcherrimus (Carl, 1909) accepted as Ktenostreptus annulipes Attems, 1909 (additional source)
Spirostreptus ruralis Carl, 1914 accepted as Isoporostreptus ruralis (Carl, 1914) (additional source)
Spirostreptus saussurei Carl, 1909 accepted as Gonoplectus saussurei (Carl, 1909) (additional source)
Spirostreptus solitarius Carl, 1909 accepted as Limnostreptus solitarius (Carl, 1909) (additional source)
Spirostreptus straminipes Carl, 1909 accepted as Gonoplectus straminipes (Carl, 1909) (additional source)
Spirostreptus zehntneri Carl, 1909 accepted as Thyropygus zehntneri (Carl, 1909) accepted as Remulopygus zehntneri (Carl, 1909) (additional source)
Stenostreptus hassleri Carl, 1917 accepted as Urostreptus hassleri (Carl, 1917) (additional source)
Sundanina albicans (Carl, 1902) (additional source)
Tectoporus ambiguus (Carl, 1941) (additional source)
Telodrepanum badaga Carl, 1932 (additional source)
Thyropygus cuisinieri Carl, 1917 (additional source)
Thyropygus eremitus Carl, 1941 accepted as Gnomognathus eremitus (Carl, 1941) (additional source)
Thyropygus helicogonus Carl, 1941 accepted as Gnomognathus helicogonus (Carl, 1941) (additional source)
Thyropygus interfectus Carl, 1941 accepted as Gnomognathus interfectus (Carl, 1941) (additional source)
Thyropygus macracanthus Carl, 1941 accepted as Gnomognathus macracanthus (Carl, 1941) (additional source)
Thyropygus minusculus discretus Carl, 1941 accepted as Gnomognathus minusculus discretus (Carl, 1941) (additional source)
Thyropygus minusculus striatus Carl, 1941 accepted as Gnomognathus minusculus striatus (Carl, 1941) (additional source)
Thyropygus minusculus suspectus Carl, 1941 accepted as Gnomognathus minusculus suspectus (Carl, 1941) (additional source)
Thyropygus minusculus vicinus Carl, 1941 accepted as Gnomognathus minusculus vicinus (Carl, 1941) (additional source)
Thyropygus nanus Carl, 1941 accepted as Gnomognathus nanus (Carl, 1941) (additional source)
Thyropygus nanus var. striatus Carl, 1941 accepted as Gnomognathus nanus (Carl, 1941) (additional source)
Thyropygus negotiosus Carl, 1941 accepted as Phyllogonostreptus negotiosus (Carl, 1941) (additional source)
Thyropygus subvalidus Carl, 1941 accepted as Carlogonus subvalidus (Carl, 1941) (additional source)
Thyropygus tricolor Carl, 1941 accepted as Gnomognathus tricolor (Carl, 1941) (additional source)
Thyropygus vicarius Carl, 1941 accepted as Gnomognathus vicarius (Carl, 1941) (additional source)
Trichogonostreptus ternetzi Carl, 1918 accepted as Trichogonostreptus (Trichogonostreptus) ternetzi Carl, 1918 (additional source)
Trichomorpha annulipes Carl, 1914 (additional source)
Trigoniulus andropygus impunctatus Carl, 1918 (additional source)
Unknown type BMNH, identified as Cryptocorypha areata (Carl, 1932)
Syntype MHNG MHNG-ARTO-14236, identified as Ktenostreptus acceptus Carl, 1941
Syntype MHNG MHNG-ARTO-14237, identified as Ktenostreptus acceptus Carl, 1941
Syntype MHNG MHNG-ARTO-14238, identified as Ktenostreptus acceptus Carl, 1941
Syntype MHNG MHNG-ARTO-14239, identified as Ktenostreptus acceptus Carl, 1941
Syntype MHNG MHNG-ARTO-14240, identified as Ktenostreptus acceptus Carl, 1941
Holotype MHNG MHNG-ARTO-14242, identified as Doratogonus annulipes Carl, 1917
Syntype MHNG MHNG-ARTO-14292, verbatimGeounit Neu-Caledonien: Mt. ..., identified as Canacophilus acutangulus Carl, 1926
Syntype MHNG MHNG-ARTO-14293, identified as Sundanina albicans (Carl, 1902)
Syntype MHNG MHNG-ARTO-14294, identified as Sholaphilus albidus Carl, 1932
Holotype MHNG MHNG-ARTO-14295, verbatimGeounit Soekaboemi, West-Jav..., identified as Tectoporus ambiguus (Carl, 1941)
Syntype MHNG MHNG-ARTO-14296, identified as Polydesmus americanus Carl, 1902
Lectotype MHNG MHNG-ARTO-14316, identified as Salpidobolus annulipes (Carl, 1912)
Syntype MHNG MHNG-ARTO-14328, identified as Spirobolellus ambiguus Carl, 1926
Nontype MHNG MHNG-ARTO-14329, identified as Spirobolellus ambiguus Carl, 1926
Holotype MHNG MHNG-ARTO-14350, identified as Batodesmus acceptus Carl, 1914
Lectotype MHNG MHNG-ARTO-14351, verbatimGeounit Bodega Central (Colo..., identified as Carlopeltis alatus (Carl, 1914)
Syntype MHNG MHNG-ARTO-14353, identified as Calymmodesmus andinus Carl, 1914
Syntype MHNG MHNG-ARTO-14354, identified as Calymmodesmus andinus Carl, 1914
Syntype MHNG MHNG-ARTO-14355, identified as Trigoniulus andropygus impunctatus Carl, 1918
Syntype MHNG MHNG-ARTO-14356, identified as Trigoniulus andropygus impunctatus Carl, 1918
Syntype MHNG MHNG-ARTO-14357, identified as Trichomorpha annulipes Carl, 1914
Syntype MHNG MHNG-ARTO-14358, identified as Trichomorpha annulipes Carl, 1914
Syntype MHNG MHNG-ARTO-14359, identified as Leptodesmus augustus Carl, 1914
Holotype MHNG MHNG-ARTO-14360, identified as Nesorthomorpha armata (Carl, 1902)
Unknown type MHNG MHNG-ARTO-14362, verbatimGeounit Palnis inf., Tandiku..., identified as Polydrepanum asperrimum (Carl, 1932)
Lectotype MHNG MHNG-ARTO-14363, verbatimGeounit Maryland, Palnis inf..., identified as Polydrepanum asperrimum (Carl, 1932)
Syntype MHNG MHNG-ARTO-14364, verbatimGeounit Travancore Vattavada..., identified as Cryptocorypha areata (Carl, 1932)
Syntype MHNG MHNG-ARTO-14365, verbatimGeounit Palnis sup. Vandarav..., identified as Cryptocorypha areata (Carl, 1932)
Syntype MHNG MHNG-ARTO-14366, verbatimGeounit Palnis sup. Kukkal-S..., identified as Cryptocorypha areata (Carl, 1932)
Syntype MHNG MHNG-ARTO-14367, identified as Cryptocorypha areata (Carl, 1932)
Syntype MHNG MHNG-ARTO-14368, verbatimGeounit Palnis sup. Maryian-..., identified as Cryptocorypha areata (Carl, 1932)
Syntype MHNG MHNG-ARTO-14369, verbatimGeounit Palnis sup. Kukkal, ..., identified as Cryptocorypha areata (Carl, 1932)
Syntype MHNG MHNG-ARTO-14370, verbatimGeounit Palnis infér. Tandik..., identified as Cryptocorypha areata (Carl, 1932)
Syntype MHNG MHNG-ARTO-14371, verbatimGeounit Palnis sup. Kodaikan..., identified as Cryptocorypha areata (Carl, 1932)
Lectotype MHNG MHNG-ARTO-14376, verbatimGeounit San Bernardino (Para..., identified as Argentocricus bernardinensis (Carl, 1918)
Lectotype MHNG MHNG-ARTO-14377, identified as Lophostreptus bicolor Carl, 1909
Paralectotype MHNG MHNG-ARTO-14378, identified as Lophostreptus bicolor Carl, 1909
Paralectotype MHNG MHNG-ARTO-14379, identified as Lophostreptus bicolor Carl, 1909
Lectotype MHNG MHNG-ARTO-14417, verbatimGeounit Njarugenje (Central-..., identified as Brachyspirobolus aequatorialis (Carl, 1909)
Unknown type MHNG MHNG-ARTO-14418, identified as Brachyspirobolus aequatorialis (Carl, 1909)
Holotype NMB NMBDIPL- 00462-Ia, identified as Prepodesmus ornatus (Peters, 1864)
Holotype NMB NMB-DIPL-00111a, verbatimGeounit Gipfel des Bowonglan..., identified as Hyleoglomeris alticola (Carl, 1912)
Paralectotype NMB NMB-DIPL-00137a, identified as Mastigorhacus alatus (Carl, 1912)
Lectotype NMB NMB-DIPL-00138a, identified as Erythrhacus arietis (Carl, 1912)
Syntype NMB NMB-DIPL-00139a, identified as Opisthoporodesmus bacillifer Carl, 1912
Syntype NMB NMB-DIPL-00162a, identified as Salpidobolus annulipes (Carl, 1912)
Paralectotype NMB NMB-DIPL-00178a, identified as Lophostreptus bicolor Carl, 1909
Syntype NMB NMB-DIPL-00222a, identified as Leptodesmus augustus Carl, 1914
Syntype NMB NMB-DIPL-00227a, identified as Trichomorpha annulipes Carl, 1914
Syntype NMB NMB-DIPL-00345a, NMB-DIPL-00345b and NMB-DIPL-00345c, identified as Agathodesmus baccatus (Carl, 1926)
Syntype NMB-DIPL-00363a to NMB-DIPL-00363g, identified as Spirobolellus ambiguus Carl, 1926
Lectotype SMF 829, identified as Mastigorhacus alatus (Carl, 1912)
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