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Hollier, J.; Stöckli, E.; Wesener, T.; Mesibov, R.; Decker, P.; Sierwald, P. (2020). An annotated list of the millipede (Diplopoda) species described by Johann Carl. Revue suisse de Zoologie. 127(1): 183-240.
381271
10.35929/RSZ.0015 [view]
Hollier, J.; Stöckli, E.; Wesener, T.; Mesibov, R.; Decker, P.; Sierwald, P.
2020
An annotated list of the millipede (Diplopoda) species described by Johann Carl
Revue suisse de Zoologie
127(1): 183-240
Publication
Johann Carl described 390 species or subspecies of millipedes during his long career at the Muséum d’histoire naturelle de Genève, studying specimens collected on his own expeditions and on those of other, mainly Swiss, naturalists. The type specimens deposited in Geneva are enumerated, and the type specimens that could be located in other collections in other museums are reported. Lectotypes are designated for 17 species: Brachyspirobolus aequatorialis (Carl, 1909), Carlopeltis alatus (Carl, 1914), Mastigorhacus alatus (Carl, 1912), Salpidobolus annulipes (Carl, 1912), Erythrhacus arietis (Carl, 1912), Argentocricus bernardinensis (Carl, 1918), Trigoniulus bitaeniatus Carl, 1912, Desmocricus conjunctus Carl, 1918, Polylepis elberti Carl, 1912, Trigoniulus incommodus Carl, 1912, Stenobolus insularis Carl, 1918, Salpidobolus lateralis (Carl, 1912), Dinematocricus lombokensis (Carl, 1912), Prepodesmus longipes (Carl, 1913), Arostrophus mertoni (Carl, 1912), Salpidobolus moenensis (Carl, 1912) and Eucarlia velox (Carl, 1912). Polydesmus carli nom. nov. is introduced as a replacement name for Polydesmus japonicus Carl, 1902, a junior homonym of Polydesmus japonicus Peters, 1864. Odontopyge johanncarli nom. nov. is introduced as a replacement name for Odontopyge fasciata Carl, 1905, a junior homonym of Odontopyge fasciata Attems, 1896.
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Acanthodesmus mortoni (Carl, 1909) (additional source)
Aceratophallus unicolor Carl, 1902 (additional source)
Acladocricus carli (Attems, 1914) (additional source)
Acladocricus macassarensis (Carl, 1912) (additional source)
Acutangulus neglectus Carl, 1903 (additional source)
Agastrophus cinctellus Carl, 1926 accepted as Hypocambala cinctella (Carl, 1926) (additional source)
Agathodesmus baccatus (Carl, 1926) (additional source)
Akreiodesmus minutus Carl, 1932 (additional source)
Akreiodesmus simulans Carl, 1932 (additional source)
Alocodesmus intermedius Carl, 1914 (additional source)
Amplinus convexus (Carl, 1902) (additional source)
Anoplodesmus cylindricus (Carl, 1935) (additional source)
Anoplodesmus humberti (Carl, 1902) (additional source)
Anoplodesmus rufocinctus (Carl, 1932) (additional source)
Anoplodesmus subcylindricus (Carl, 1932) (additional source)
Antheromorpha mediovirgata (Carl, 1941) (additional source)
Antheromorpha orophila (Carl, 1941) (additional source)
Antichirogonus hirta (Carl, 1932) (additional source)
Antichirogonus laevisulcata (Carl, 1932) (additional source)
Aphelidesmus ambiguus Carl, 1914 (additional source)
Aporodesmus sellae Silvestri, 1907 (additional source)
Argentocricus bernardinensis (Carl, 1918) (additional source)
Arostrophus mertoni (Carl, 1912) (additional source)
Atopochetus sumatranus (Carl, 1906) (additional source)
Aulacobolus dysoni Carl, 1941 (additional source)
Aulacobolus ejaculans Carl, 1941 (additional source)
Aulacobolus ejaculans vallensis Carl, 1941 (additional source)
Aulacobolus gravelyi Silvestri, 1916 (additional source)
Aulacobolus perfidus Carl, 1941 (additional source)
Aulodesmus flavosignatus (Carl, 1909) (additional source)
Barydesmus propinquus (Carl, 1902) (additional source)
Barydesmus riparius (Carl, 1902) (additional source)
Batodesmus acceptus Carl, 1914 (additional source)
Brachycerodesmus fuhrmanni Carl, 1914 (additional source)
Brachycerodesmus petersi Carl, 1914 (additional source)
Brachyspirobolus aequatorialis (Carl, 1909) accepted as Microspirobolus aequatorialis Carl, 1909 accepted as Bukobolus aequatorialis (Carl, 1909) (additional source)
Brachyspirobolus domesticus (Carl, 1909) (additional source)
Bucinogonus kandti (Carl, 1909) (additional source)
Calymmodesmus andinus Carl, 1914 (additional source)
Campodesmus costatus (Carl, 1913) (additional source)
Canacophilus acutangulus Carl, 1926 (additional source)
Canacophilus fastidiosus Carl, 1926 (additional source)
Canacophilus humboldti Carl, 1926 (additional source)
Canacophilus lifouensis Carl, 1926 (additional source)
Canacophilus rouxi Carl, 1926 (additional source)
Canacophilus sarasini Carl, 1926 (additional source)
Carlocricus elberti (Carl, 1912) (additional source)
Carlogonus subvalidus (Carl, 1941) (additional source)
Carlopeltis alatus (Carl, 1914) (additional source)
Carlotretus setosus (Carl, 1922) (additional source)
Castanotherium boetonense Carl, 1912 (additional source)
Castanotherium cinctum (Carl, 1906) (additional source)
Castanotherium decoratum Carl, 1912 (additional source)
Castanotherium distinctum Carl, 1912 (additional source)
Castanotherium laeve Carl, 1912 (additional source)
Castanotherium ornatum Carl, 1912 (additional source)
Castanotherium pilosum Carl, 1912 (additional source)
Castanotherium simplex (Carl, 1909) (additional source)
Castanotherium sparsepunctatum Carl, 1912 (additional source)
Castanotherium stellatum Carl, 1912 (additional source)
Castanotherium suspectum Carl, 1912 (additional source)
Catharosoma intermedium (Carl, 1902) (additional source)
Centrelus neglectus (Carl, 1919) (additional source)
Chondrodesmus attemsi Carl, 1914 (additional source)
Chondrodesmus carbonarius Carl, 1914 (additional source)
Chondrodesmus convexus Carl, 1914 (additional source)
Chondrodesmus dorsovittatus Carl, 1914 (additional source)
Chondrodesmus granosus (Carl, 1902) (additional source)
Chondrodesmus nobilis Carl, 1914 (additional source)
Chondrodesmus riparius Carl, 1914 (additional source)
Chondromorpha kelaarti valparaiensis (Carl, 1932) accepted as Chondromorpha kelaarti (Humbert, 1865) (additional source)
Cingalobolus bugnioni Carl, 1918 (additional source)
Columbianum gracilicorne (Carl, 1914) (additional source)
Coonoorophilus monstruosus Carl, 1932 (additional source)
Coromus lugubris (Carl, 1909) (additional source)
Cotylotropis hystrix Carl, 1926 (additional source)
Cryptocorypha areata (Carl, 1932) (additional source)
Cryptocorypha kandyanus (Carl, 1932) (additional source)
Cryptocorypha riparia (Carl, 1932) (additional source)
Cryptocorypha tuberculata (Carl, 1932) (additional source)
Cryptodesmus triseriatus Carl, 1912 (additional source)
Cyrtodesmus ammonites (Carl, 1914) (additional source)
Cyrtodesmus hirsutus (Carl, 1914) (additional source)
Cyrtodesmus niger (Carl, 1914) (additional source)
Delarthrum hingstoni (Carl, 1935) (additional source)
Delarthrum rugulosum (Carl, 1932) (additional source)
Delarthrum simulans (Carl, 1935) (additional source)
Desmocricus conjunctus Carl, 1918 (additional source)
Desmoxytes bicolor (Carl, 1902) (additional source)
Dinematocricus faucium Brölemann, 1913 accepted as Salpidobolus faucium (Brölemann, 1913) (additional source)
Dinematocricus leucopygus (Carl, 1912) accepted as Salpidobolus leucopygus (Carl, 1912) (additional source)
Dinematocricus lombokensis (Carl, 1912) accepted as Salpidobolus lombokensis (Carl, 1912) (additional source)
Diopsiulus greeni Carl, 1941 (additional source)
Doratogonus annulipes Carl, 1917 (additional source)
Enghoffosoma spinipleurum (Carl, 1941) (additional source)
Epinannolene exilis Carl, 1914 (additional source)
Epinannolene fuhrmanni Carl, 1914 (additional source)
Epinannolene nigrita Carl, 1914 (additional source)
Erythrhacus arietis (Carl, 1912) (additional source)
Erythrhacus sarasinorum (Carl, 1912) (additional source)
Unknown type BMNH, identified as Cryptocorypha areata (Carl, 1932)
Syntype SMF, identified as Trigoniulus bitaeniatus Carl, 1912
Syntype ZMHB, identified as Holopodostreptus braueri Carl, 1913
Syntype ZMHB, identified as Sculptoteles braueri (Carl, 1918)
Holotype ZMHB, verbatimGeounit Tjibodas, Zentraljav..., identified as Orthomorpha butteli (Carl, 1922)
Syntype ZMUH, identified as Odontopyge dorsalis Carl, 1909
Syntype ZMUH, identified as Microspirobolus domesticus Carl, 1909
Syntype BMNH, identified as Hingstonia eremita Carl, 1935
Paralectotype ZMHB, identified as Platyrhacus fecundus Carl, 1912
Syntype ZMUH, identified as Euryzonus flavosignatus Carl, 1909
Syntype MCZL, identified as Euryzonus flavosignatus Carl, 1909
Syntype ZMHB, identified as Euryzonus flavosignatus Carl, 1909
Syntype ZMUH, identified as Strongylosoma fossiger var. typica Carl, 1909
Syntype ZMHB, identified as Strongylosoma fossiger var. typica Carl, 1909
Syntype MCZL, identified as Strongylosoma fossiger var. typica Carl, 1909
Syntype ZMUH, identified as Strongylosoma fossiger silvestre Carl, 1909
Syntype MCZL, identified as Strongylosoma fossiger silvestre Carl, 1909
Syntype ZMUH, identified as Eviulisoma fossiger ussuwiense (Carl, 1909)
Syntype MCZL, identified as Eviulisoma fossiger ussuwiense (Carl, 1909)
Syntype MCZL, identified as Gonoplectus frater (Carl, 1909)
Holotype ZMHB, identified as Siphonophora grandis Carl, 1922
Syntype BMNH, identified as Aulacobolus gravelyi Silvestri, 1916
Holotype ZMHB, identified as Pycnotropis haenschi Carl, 1918
Syntype BMNH, identified as Xiphidiogonus hendersoni Carl, 1932
Unknown type BMNH, identified as Orthomorpha hingstoni Carl, 1935
Holotype BMNH, identified as Trachyiulus humberti willeyi Carl, 1941
Holotype BMNH, identified as Telodrepanum implicatum (Carl, 1941)
Syntype ZMHB, identified as Rhinocricus instabilis adolescens Carl, 1914
Syntype ZMHB, identified as Rhinocricus instabilis valens Carl, 1914
Syntype ZMUH, identified as Odontopyge intermedia Carl, 1909
Syntype ZMUH, identified as Bucinogonus kandti (Carl, 1909)
Syntype MCZL, identified as Bucinogonus kandti (Carl, 1909)
Syntype ZMHB, identified as Bucinogonus kandti (Carl, 1909)
Syntype SMF, verbatimGeounit Celebes: Roembi-Meng..., identified as Rhinocricus lateralis Carl, 1912
Unknown type SMF, identified as Dinematocricus lombokensis (Carl, 1912)
Syntype ZMUH, identified as Morphotelus mareesi (Carl, 1909)
Syntype MCZL, identified as Morphotelus mareesi (Carl, 1909)
Syntype ZMHB, identified as Morphotelus mareesi (Carl, 1909)
Holotype BMNH, verbatimGeounit Nördliche Chin-Hills..., identified as Antheromorpha mediovirgata (Carl, 1941)
Holotype ZMHB, verbatimGeounit Maxwells Hill (Distr..., identified as Siphonophora minuta Carl, 1922
Unknown type BMNH, identified as Substrongylosoma montigena (Carl, 1935)
Holotype BMNH, identified as Phyllogonostreptus negotiosus (Carl, 1941)
Syntype ZMUH, identified as Helicochetus pococki (Carl, 1909)
Syntype ZMUH, identified as Odontopyge regina Carl, 1909
Syntype BMNH, identified as Orthomorpha simulans Carl, 1935
Paralectotype ZMUH, identified as Odontopyge socialis Carl, 1909
Paralectotype MCZL, identified as Odontopyge socialis Carl, 1909
Syntype ZMUH, identified as Odontopyge spiralis Carl, 1909
Unknown type MCZL, identified as Thyropygus straminipes (Carl, 1909)
Syntype BMNH, identified as Polydrepanum tamilum Carl, 1932
Syntype ZMUH, identified as Strongylosoma tesselatum Carl, 1909
Syntype MCZL, identified as Strongylosoma tesselatum Carl, 1909
Holotype BMNH, identified as Sundanina trifida Carl, 1941
Syntype ZMUH, identified as Strongylosoma vagans Carl, 1909
Syntype MCZL, identified as Strongylosoma vagans Carl, 1909
Syntype ZMHB, identified as Strongylosoma vagans Carl, 1909
Syntype MHNG (MHNG-ARTO-14342, verbatimGeounit Tschalabel N.-C. unt..., identified as Spirobolellus sarasini Carl, 1926
Syntype MHNG MHNGARTO- 14250, identified as Steganostigmus contortipes Carl, 1932
Syntype MHNG MHNG-ARTO- 14480, identified as Diopsiulus vagans Carl, 1941
Syntype MHNG MHNG-ARTO-14229, verbatimGeounit Costarica, St. José,..., identified as Platyrhacus fraternus Carl, 1902
Syntype MHNG MHNG-ARTO-14230, identified as Trichozonus escalerae Carl, 1905
Holotype MHNG MHNG-ARTO-14231, identified as Pagodesmus eremitus Carl, 1932
Holotype MHNG MHNG-ARTO-14232, identified as Podoglyphiulus errabundus (Carl, 1941)
Holotype MHNG MHNG-ARTO-14233, identified as Gnomognathus eremitus (Carl, 1941)
Syntype MHNG MHNG-ARTO-14234, identified as Picrogonopus gracillimus Carl, 1941
Syntype MHNG MHNG-ARTO-14235, identified as Picrogonopus gracillimus Carl, 1941
Syntype MHNG MHNG-ARTO-14236, identified as Ktenostreptus acceptus Carl, 1941
Syntype MHNG MHNG-ARTO-14237, identified as Ktenostreptus acceptus Carl, 1941
Syntype MHNG MHNG-ARTO-14238, identified as Ktenostreptus acceptus Carl, 1941
Syntype MHNG MHNG-ARTO-14239, identified as Ktenostreptus acceptus Carl, 1941
Syntype MHNG MHNG-ARTO-14240, identified as Ktenostreptus acceptus Carl, 1941
Holotype MHNG MHNG-ARTO-14242, identified as Doratogonus annulipes Carl, 1917
Syntype MHNG MHNG-ARTO-14243, identified as Pagodesmus biporus Carl, 1932
Syntype MHNG MHNG-ARTO-14244, verbatimGeounit Palnis, Pumbarai and..., identified as Pagodesmus biporus Carl, 1932
Syntype MHNG MHNG-ARTO-14245, identified as Pagodesmus biporus Carl, 1932
Holotype MHNG MHNG-ARTO-14246, identified as Steganostigmus canonicus Carl, 1932
Holotype MHNG MHNG-ARTO-14247, identified as Pseudosphaeroparia cardamomi Carl, 1932
Paratype MHNG MHNG-ARTO-14248, identified as Pseudosphaeroparia cardamomi Carl, 1932
Syntype MHNG MHNG-ARTO-14249, identified as Steganostigmus contortipes Carl, 1932
Holotype MHNG MHNG-ARTO-14251, verbatimGeounit "Elizabethville (Kat..., identified as Poratophilus distinctus Carl, 1917
Holotype MHNG MHNG-ARTO-14252, identified as Xiphidiogonus dravidus Carl, 1932
Lectotype MHNG MHNG-ARTO-14253, identified as Stenostreptus hassleri Carl, 1917
Paralectotype MHNG MHNG-ARTO-14254 and MHNG-ARTO-14255, identified as Urostreptus hassleri (Carl, 1917)
Syntype MHNG MHNG-ARTO-14256, verbatimGeounit Kleiner Djungel 24.X..., identified as Thyropygus helicogonus Carl, 1941
Lectotype MHNG MHNG-ARTO-14258, identified as Poratia heterotuberculata Carl, 1902
Syntype MHNG MHNG-ARTO-14259, identified as Poratia heterotuberculata Carl, 1902
Paralectotype MHNG MHNG-ARTO-14259a, identified as Poratia heterotuberculata Carl, 1902
Syntype MHNG MHNG-ARTO-14260, identified as Poratia heterotuberculata Carl, 1902
Holotype MHNG MHNG-ARTO-14261, verbatimGeounit Travancore, Vattavad..., identified as Sundanina hirta Carl, 1932
Syntype MHNG MHNG-ARTO-14262, identified as Trachyjulus humberti Carl, 1911
Lectotype MHNG MHNG-ARTO-14263, identified as Stenobolus insularis Carl, 1918
Unknown type MHNG MHNG-ARTO-14264, identified as Stenobolus insularis Carl, 1918
Unknown type MHNG MHNG-ARTO-14265, verbatimGeounit Male Atoll, Maledive..., identified as Stenobolus insularis Carl, 1918
Holotype MHNG MHNG-ARTO-14266, identified as Thyropygus interfectus Carl, 1941
Syntype MHNG MHNG-ARTO-14267, identified as Glyphiulus javanicus Carl, 1911
Syntype MHNG MHNG-ARTO-14268, verbatimGeounit Shilowana, Afrique m..., identified as Zinophora junodi (Carl, 1917)
Syntype MHNG MHNG-ARTO-14269, verbatimGeounit Ricalla, Afrique mér..., identified as Zinophora junodi (Carl, 1917)
Syntype MHNG MHNG-ARTO-14270, identified as Chondromorpha kelaarti valparaiensis (Carl, 1932)
Lectotype MHNG MHNG-ARTO-14271, verbatimGeounit Palnis, Vandavravu s..., identified as Antichirogonus laevisulcata (Carl, 1932)
Paralectotype MHNG MHNG-ARTO-14272, identified as Antichirogonus laevisulcata (Carl, 1932)
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