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Hollier, J.; Stöckli, E.; Wesener, T.; Mesibov, R.; Decker, P.; Sierwald, P. (2020). An annotated list of the millipede (Diplopoda) species described by Johann Carl. Revue suisse de Zoologie. 127(1): 183-240.
381271
10.35929/RSZ.0015 [view]
Hollier, J.; Stöckli, E.; Wesener, T.; Mesibov, R.; Decker, P.; Sierwald, P.
2020
An annotated list of the millipede (Diplopoda) species described by Johann Carl
Revue suisse de Zoologie
127(1): 183-240
Publication
Johann Carl described 390 species or subspecies of millipedes during his long career at the Muséum d’histoire naturelle de Genève, studying specimens collected on his own expeditions and on those of other, mainly Swiss, naturalists. The type specimens deposited in Geneva are enumerated, and the type specimens that could be located in other collections in other museums are reported. Lectotypes are designated for 17 species: Brachyspirobolus aequatorialis (Carl, 1909), Carlopeltis alatus (Carl, 1914), Mastigorhacus alatus (Carl, 1912), Salpidobolus annulipes (Carl, 1912), Erythrhacus arietis (Carl, 1912), Argentocricus bernardinensis (Carl, 1918), Trigoniulus bitaeniatus Carl, 1912, Desmocricus conjunctus Carl, 1918, Polylepis elberti Carl, 1912, Trigoniulus incommodus Carl, 1912, Stenobolus insularis Carl, 1918, Salpidobolus lateralis (Carl, 1912), Dinematocricus lombokensis (Carl, 1912), Prepodesmus longipes (Carl, 1913), Arostrophus mertoni (Carl, 1912), Salpidobolus moenensis (Carl, 1912) and Eucarlia velox (Carl, 1912). Polydesmus carli nom. nov. is introduced as a replacement name for Polydesmus japonicus Carl, 1902, a junior homonym of Polydesmus japonicus Peters, 1864. Odontopyge johanncarli nom. nov. is introduced as a replacement name for Odontopyge fasciata Carl, 1905, a junior homonym of Odontopyge fasciata Attems, 1896.
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Agastrophus cinctellus Carl, 1926 accepted as Hypocambala cinctella (Carl, 1926) (additional source)
Agathodesmus baccatus (Carl, 1926) (additional source)
Alocodesmus intermedius Carl, 1914 (additional source)
Amplinus convexus (Carl, 1902) (additional source)
Anoplodesmus cylindricus (Carl, 1935) (additional source)
Anoplodesmus humberti (Carl, 1902) (additional source)
Antichirogonus hirta (Carl, 1932) (additional source)
Antichirogonus laevisulcata (Carl, 1932) (additional source)
Aphelidesmus ambiguus Carl, 1914 (additional source)
Argentocricus bernardinensis (Carl, 1918) (additional source)
Aulacobolus dysoni Carl, 1941 (additional source)
Aulacobolus ejaculans Carl, 1941 (additional source)
Aulacobolus ejaculans vallensis Carl, 1941 (additional source)
Aulacobolus gravelyi Silvestri, 1916 (additional source)
Aulodesmus flavosignatus (Carl, 1909) (additional source)
Batodesmus acceptus Carl, 1914 (additional source)
Brachycerodesmus fuhrmanni Carl, 1914 (additional source)
Brachyspirobolus aequatorialis (Carl, 1909) (additional source)
Brachyspirobolus domesticus (Carl, 1909) (additional source)
Bucinogonus kandti (Carl, 1909) (additional source)
Calymmodesmus andinus Carl, 1914 (additional source)
Campodesmus costatus (Carl, 1913) (additional source)
Canacophilus acutangulus Carl, 1926 (additional source)
Canacophilus fastidiosus Carl, 1926 (additional source)
Canacophilus humboldti Carl, 1926 (additional source)
Carlocricus elberti (Carl, 1912) (additional source)
Carlopeltis alatus (Carl, 1914) (additional source)
Castanotherium boetonense Carl, 1912 (additional source)
Castanotherium cinctum (Carl, 1906) (additional source)
Castanotherium decoratum Carl, 1912 (additional source)
Castanotherium distinctum Carl, 1912 (additional source)
Castanotherium laeve Carl, 1912 (additional source)
Catharosoma intermedium (Carl, 1902) (additional source)
Chondrodesmus attemsi Carl, 1914 (additional source)
Chondrodesmus carbonarius Carl, 1914 (additional source)
Chondrodesmus convexus Carl, 1914 (additional source)
Chondrodesmus dorsovittatus Carl, 1914 (additional source)
Chondromorpha kelaarti valparaiensis (Carl, 1932) accepted as Chondromorpha kelaarti (Humbert, 1865) (additional source)
Cingalobolus bugnioni Carl, 1918 (additional source)
Columbianum gracilicorne (Carl, 1914) (additional source)
Cotylotropis hystrix Carl, 1926 (additional source)
Cryptocorypha areata (Carl, 1932) (additional source)
Cryptocorypha kandyanus (Carl, 1932) (additional source)
Cyrtodesmus ammonites (Carl, 1914) (additional source)
Cyrtodesmus hirsutus (Carl, 1914) (additional source)
Delarthrum hingstoni (Carl, 1935) (additional source)
Desmocricus conjunctus Carl, 1918 (additional source)
Desmoxytes bicolor (Carl, 1902) (additional source)
Dinematocricus faucium Brölemann, 1913 accepted as Salpidobolus faucium (Brölemann, 1913) (additional source)
Diopsiulus greeni Carl, 1941 (additional source)
Doratogonus annulipes Carl, 1917 (additional source)
Epinannolene exilis Carl, 1914 (additional source)
Epinannolene fuhrmanni Carl, 1914 (additional source)
Erythrhacus arietis (Carl, 1912) (additional source)
Eviulisoma fossiger (Carl, 1909) (additional source)
Eviulisoma fossiger ussuwiense (Carl, 1909) accepted as Strongylosoma fossiger Carl, 1909 (additional source)
Eviulisoma silvestre (Carl, 1909) (additional source)
Geotypodon dispersus (Carl, 1909) (additional source)
Geotypodon intermedius (Carl, 1909) (additional source)
Glomeridesmus indus Carl, 1942 (additional source)
Glyphiulus javanicus Carl, 1911 (additional source)
Gnomognathus eremitus (Carl, 1941) (additional source)
Gnomognathus helicogonus (Carl, 1941) (additional source)
Gnomognathus interfectus (Carl, 1941) (additional source)
Gonoplectus frater (Carl, 1909) (additional source)
Gyrophallus imitans Carl, 1914 (additional source)
Haplothysanus dorsalis (Carl, 1909) (additional source)
Haplothysanus emini (Carl, 1909) (additional source)
Harpagomorpha dentata (Carl, 1932) (additional source)
Harpurostreptus matarae Carl, 1941 (additional source)
Harpurostreptus montivagus Carl, 1941 (additional source)
Hingstonia eremita Carl, 1935 (additional source)
Holopodostreptus braueri Carl, 1913 (additional source)
Hoplurorhachis crassipes (Carl, 1909) (additional source)
Hyleoglomeris alticola (Carl, 1912) (additional source)
Hyleoglomeris eremita (Carl, 1912) (additional source)
Hyloecostreptus inconstans (Carl, 1914) (additional source)
Hypocambala caledonica (Carl, 1926) (additional source)
Klimakodesmus gravelyi Carl, 1932 (additional source)
Ktenostreptus acceptus Carl, 1941 (additional source)
Kukkalodesmus exiguus Carl, 1932 (additional source)
Lankabolus coelebs Carl, 1941 accepted as Lankabolus greeni (Pocock, 1892) (additional source)
Leptodesmus augustus Carl, 1914 (additional source)
Leptostreptus caudatus Carl, 1941 accepted as Leptostreptus caudiculatus (Karsch, 1881) (additional source)
Lophostreptus bicolor Carl, 1909 (additional source)
Lophostreptus ptilostreptoides Carl, 1909 (additional source)
Mastigorhacus alatus (Carl, 1912) (additional source)
Microspirobolus aequatorialis Carl, 1909 accepted as Bukobolus aequatorialis (Carl, 1909) accepted as Brachyspirobolus aequatorialis (Carl, 1909) (additional source)
Neocordyloporus asperus Carl, 1905 (additional source)
Neodontopyge gracilis (Carl, 1913) (additional source)
Nesorthomorpha armata (Carl, 1902) (additional source)
Nesorthomorpha coriacea (Carl, 1902) (additional source)
Nyssodesmus bivirgatus (Carl, 1902) accepted as Nyssodesmus phyton (Peters, 1864) (additional source)
Nyssodesmus fraternus (Carl, 1902) (additional source)
Odontopyge johanncarli Hollier, Stöckli, Wesener, Mesibov, Decker & Sierwald, 2020 (status source)
Odontopyge kandti Carl, 1909 (additional source)
Oniscodesmus fuhrmanni Carl, 1914 (additional source)
Ootacadesmus humilis Carl, 1932 (additional source)
Opisthoporodesmus bacillifer Carl, 1912 (additional source)
Orthomorpha butteli (Carl, 1922) (additional source)
Unknown type BMNH, identified as Cryptocorypha areata (Carl, 1932)
Syntype SMF, identified as Trigoniulus bitaeniatus Carl, 1912
Syntype ZMHB, identified as Holopodostreptus braueri Carl, 1913
Syntype ZMHB, identified as Sculptoteles braueri (Carl, 1918)
Holotype ZMHB, verbatimGeounit Tjibodas, Zentraljav..., identified as Orthomorpha butteli (Carl, 1922)
Syntype ZMUH, identified as Odontopyge dorsalis Carl, 1909
Syntype ZMUH, identified as Microspirobolus domesticus Carl, 1909
Syntype BMNH, identified as Hingstonia eremita Carl, 1935
Paralectotype ZMHB, identified as Platyrhacus fecundus Carl, 1912
Syntype ZMUH, identified as Euryzonus flavosignatus Carl, 1909
Syntype MCZL, identified as Euryzonus flavosignatus Carl, 1909
Syntype ZMHB, identified as Euryzonus flavosignatus Carl, 1909
Syntype ZMUH, identified as Strongylosoma fossiger var. typica Carl, 1909
Syntype ZMHB, identified as Strongylosoma fossiger var. typica Carl, 1909
Syntype MCZL, identified as Strongylosoma fossiger var. typica Carl, 1909
Syntype ZMUH, identified as Strongylosoma fossiger silvestre Carl, 1909
Syntype MCZL, identified as Strongylosoma fossiger silvestre Carl, 1909
Syntype ZMUH, identified as Eviulisoma fossiger ussuwiense (Carl, 1909)
Syntype MCZL, identified as Eviulisoma fossiger ussuwiense (Carl, 1909)
Syntype MCZL, identified as Gonoplectus frater (Carl, 1909)
Holotype ZMHB, identified as Siphonophora grandis Carl, 1922
Syntype BMNH, identified as Aulacobolus gravelyi Silvestri, 1916
Holotype ZMHB, identified as Pycnotropis haenschi Carl, 1918
Syntype BMNH, identified as Xiphidiogonus hendersoni Carl, 1932
Unknown type BMNH, identified as Orthomorpha hingstoni Carl, 1935
Holotype BMNH, identified as Trachyiulus humberti willeyi Carl, 1941
Holotype BMNH, identified as Telodrepanum implicatum (Carl, 1941)
Syntype ZMHB, identified as Rhinocricus instabilis adolescens Carl, 1914
Syntype ZMHB, identified as Rhinocricus instabilis valens Carl, 1914
Syntype ZMUH, identified as Odontopyge intermedia Carl, 1909
Syntype ZMUH, identified as Bucinogonus kandti (Carl, 1909)
Syntype MCZL, identified as Bucinogonus kandti (Carl, 1909)
Syntype ZMHB, identified as Bucinogonus kandti (Carl, 1909)
Syntype SMF, verbatimGeounit Celebes: Roembi-Meng..., identified as Rhinocricus lateralis Carl, 1912
Syntype MHNG MHNGARTO- 14250, identified as Steganostigmus contortipes Carl, 1932
Syntype MHNG MHNG-ARTO-14229, verbatimGeounit Costarica, St. José,..., identified as Platyrhacus fraternus Carl, 1902
Syntype MHNG MHNG-ARTO-14230, identified as Trichozonus escalerae Carl, 1905
Holotype MHNG MHNG-ARTO-14231, identified as Pagodesmus eremitus Carl, 1932
Holotype MHNG MHNG-ARTO-14232, identified as Podoglyphiulus errabundus (Carl, 1941)
Holotype MHNG MHNG-ARTO-14233, identified as Gnomognathus eremitus (Carl, 1941)
Syntype MHNG MHNG-ARTO-14234, identified as Picrogonopus gracillimus Carl, 1941
Syntype MHNG MHNG-ARTO-14235, identified as Picrogonopus gracillimus Carl, 1941
Syntype MHNG MHNG-ARTO-14236, identified as Ktenostreptus acceptus Carl, 1941
Syntype MHNG MHNG-ARTO-14237, identified as Ktenostreptus acceptus Carl, 1941
Syntype MHNG MHNG-ARTO-14238, identified as Ktenostreptus acceptus Carl, 1941
Syntype MHNG MHNG-ARTO-14239, identified as Ktenostreptus acceptus Carl, 1941
Syntype MHNG MHNG-ARTO-14240, identified as Ktenostreptus acceptus Carl, 1941
Holotype MHNG MHNG-ARTO-14242, identified as Doratogonus annulipes Carl, 1917
Syntype MHNG MHNG-ARTO-14243, identified as Pagodesmus biporus Carl, 1932
Syntype MHNG MHNG-ARTO-14244, verbatimGeounit Palnis, Pumbarai and..., identified as Pagodesmus biporus Carl, 1932
Syntype MHNG MHNG-ARTO-14245, identified as Pagodesmus biporus Carl, 1932
Holotype MHNG MHNG-ARTO-14246, identified as Steganostigmus canonicus Carl, 1932
Holotype MHNG MHNG-ARTO-14247, identified as Pseudosphaeroparia cardamomi Carl, 1932
Paratype MHNG MHNG-ARTO-14248, identified as Pseudosphaeroparia cardamomi Carl, 1932
Syntype MHNG MHNG-ARTO-14249, identified as Steganostigmus contortipes Carl, 1932
Holotype MHNG MHNG-ARTO-14251, verbatimGeounit "Elizabethville (Kat..., identified as Poratophilus distinctus Carl, 1917
Holotype MHNG MHNG-ARTO-14252, identified as Xiphidiogonus dravidus Carl, 1932
Lectotype MHNG MHNG-ARTO-14253, identified as Stenostreptus hassleri Carl, 1917
Paralectotype MHNG MHNG-ARTO-14254 and MHNG-ARTO-14255, identified as Urostreptus hassleri (Carl, 1917)
Syntype MHNG MHNG-ARTO-14256, verbatimGeounit Kleiner Djungel 24.X..., identified as Thyropygus helicogonus Carl, 1941
Lectotype MHNG MHNG-ARTO-14258, identified as Poratia heterotuberculata Carl, 1902
Syntype MHNG MHNG-ARTO-14259, identified as Poratia heterotuberculata Carl, 1902
Paralectotype MHNG MHNG-ARTO-14259a, identified as Poratia heterotuberculata Carl, 1902
Syntype MHNG MHNG-ARTO-14260, identified as Poratia heterotuberculata Carl, 1902
Holotype MHNG MHNG-ARTO-14261, verbatimGeounit Travancore, Vattavad..., identified as Sundanina hirta Carl, 1932
Syntype MHNG MHNG-ARTO-14262, identified as Trachyjulus humberti Carl, 1911
Lectotype MHNG MHNG-ARTO-14263, identified as Stenobolus insularis Carl, 1918
Unknown type MHNG MHNG-ARTO-14264, identified as Stenobolus insularis Carl, 1918
Unknown type MHNG MHNG-ARTO-14265, verbatimGeounit Male Atoll, Maledive..., identified as Stenobolus insularis Carl, 1918
Holotype MHNG MHNG-ARTO-14266, identified as Thyropygus interfectus Carl, 1941
Syntype MHNG MHNG-ARTO-14267, identified as Glyphiulus javanicus Carl, 1911
Syntype MHNG MHNG-ARTO-14268, verbatimGeounit Shilowana, Afrique m..., identified as Zinophora junodi (Carl, 1917)
Syntype MHNG MHNG-ARTO-14269, verbatimGeounit Ricalla, Afrique mér..., identified as Zinophora junodi (Carl, 1917)
Syntype MHNG MHNG-ARTO-14270, identified as Chondromorpha kelaarti valparaiensis (Carl, 1932)
Lectotype MHNG MHNG-ARTO-14271, verbatimGeounit Palnis, Vandavravu s..., identified as Antichirogonus laevisulcata (Carl, 1932)
Paralectotype MHNG MHNG-ARTO-14272, identified as Antichirogonus laevisulcata (Carl, 1932)
Paralectotype MHNG MHNG-ARTO-14273, identified as Antichirogonus laevisulcata (Carl, 1932)
Syntype MHNG MHNG-ARTO-14292, verbatimGeounit Neu-Caledonien: Mt. ..., identified as Canacophilus acutangulus Carl, 1926
Syntype MHNG MHNG-ARTO-14293, identified as Sundanina albicans (Carl, 1902)
Syntype MHNG MHNG-ARTO-14294, identified as Sholaphilus albidus Carl, 1932
Holotype MHNG MHNG-ARTO-14295, verbatimGeounit Soekaboemi, West-Jav..., identified as Tectoporus ambiguus (Carl, 1941)
Syntype MHNG MHNG-ARTO-14296, identified as Polydesmus americanus Carl, 1902
Syntype MHNG MHNG-ARTO-14305, verbatimGeounit Loka Celebes mérid. ..., identified as Strongylosoma hirtipes Carl, 1912
Syntype MHNG MHNG-ARTO-14312, identified as Salpidobolus centralis (Carl, 1912)
Syntype MHNG MHNG-ARTO-14313, verbatimGeounit Manipi Celebes mérid..., identified as Rhinocricus fulvotaeniatus Carl, 1912
Syntype MHNG MHNG-ARTO-14315, identified as Rhinocricus gorontalensis Carl, 1912
Lectotype MHNG MHNG-ARTO-14316, identified as Salpidobolus annulipes (Carl, 1912)
Syntype MHNG MHNG-ARTO-14321, identified as Strongylosoma intermedium Carl, 1902
Syntype MHNG MHNG-ARTO-14322, verbatimGeounit Tchalabel N. Caled. ..., identified as Plethodesmus biseriatus Carl, 1926
Syntype MHNG MHNG-ARTO-14323, identified as Spirobolellus dimidiatus Carl, 1926
Syntype MHNG MHNG-ARTO-14324, identified as Spirobolellus fallax Carl, 1926
Syntype MHNG MHNG-ARTO-14327, verbatimGeounit Mt Canala Nlle. Cale..., identified as Spirobolellus canalensis Carl, 1926
Syntype MHNG MHNG-ARTO-14328, identified as Spirobolellus ambiguus Carl, 1926
Nontype MHNG MHNG-ARTO-14329, identified as Spirobolellus ambiguus Carl, 1926
Syntype MHNG MHNG-ARTO-14330, identified as Spirobolellus dispersus Carl, 1926
Syntype MHNG MHNG-ARTO-14331, verbatimGeounit N. Caled., Naltho, L..., identified as Spirobolellus dispersus Carl, 1926
Syntype MHNG MHNG-ARTO-14337, identified as Spirobolellus comicus Carl, 1926
Syntype MHNG MHNG-ARTO-14338, identified as Spirobolellus expulsus Carl, 1926
Syntype MHNG MHNG-ARTO-14343, identified as Rhinotus ducalis Carl, 1926
Syntype MHNG MHNG-ARTO-14345, verbatimGeounit Loyalty-Inseln: Ouvé..., identified as Rhinotus hispidus Carl, 1926
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