original description
Cook, O. F. (1896). A new diplopod fauna in Liberia. American Naturalist, 30: 413-420, available online at https://www.biodiversitylibrary.org/page/41431949, https://www.journals.uchicago.edu/doi/pdf/10.1086/276402
page(s): 419 [details]
original description
(of Euhercodesmus Schubart, 1955) Schubart, O. (1955). Proterospermophora oder Polydesmoidea von Französisch West-Afrika (Diplopoda). Bulletin de l'Institut française d'Afrique Noire, série A, 17(2): 377-443
page(s): 436 [details]
original description
(of Hercodesmus Cook, 1896) Cook, O. F. (1896). A new diplopod fauna in Liberia. American Naturalist, 30: 413-420, available online at https://www.biodiversitylibrary.org/page/41431949, https://www.journals.uchicago.edu/doi/pdf/10.1086/276402
page(s): 419 [details]
additional source
Golovatch, S. I.; Nzoko Fiemapong, A. R.; VandenSpiegel, D. (2015). Notes on Afrotropical Pyrgodesmidae, 2 (Diplopoda: Polydesmida). <em>Arthropoda Selecta.</em> 24(4): 387–400., available online at https://www.kmkjournals.com/upload/PDF/ArthropodaSelecta/24/24_4_387_400_Golovatch_et_al_for_Inet.pdf [details]
additional source
Golovatch, Sergei Ilyich; VandenSpiegel, Didier. (2014). Notes on Afrotropical Pyrgodesmidae, 1 (Diplopoda: Polydesmida). <em>Arthropoda Selecta.</em> 23 (4): 319-335., available online at https://kmkjournals.com/upload/PDF/ArthropodaSelecta/23/23_4_319_335_Golovatch_VandenSpiegel_for_Inet.pdf
note: Includes key to species in the genus on page 334; does not include U. camerunensis. [details] Available for editors 
From editor or global species database
Diagnosis A genus of Pyrgodesmidae with 19 or 20 body segments. Metatergal microsculpture typical of Pyrgodesmidae, with microvilli supporting a cerategument. Head roundish, with vertigial granulations; genae with a deep groove for accommodation of antennae. Antennae short, clavate, antennomere 5 larger than 6th, each with an apicodorsal group of bacilliform sensilla. Paraterga set low (at about lower 1/3 body height). Collum flabellate, often capuchin-shaped, entirely covering the head from above, always with clearly differentiated lobulations at anterior margin, centrally often with differentiated tubercles as well. Postcollum metaterga with usual larger mid-dorsal and dorsolateral tubercles in 2+2 paramedian crests, each crest with 2–3 transverse rows of evident to poorly expressed tubercles; paraterga 2 trilobate, following ones bilobate at lateral margin; anterolateral lobulations wanting, but a few caudolaterals sometimes present; all paraterga strongly declined ventolaterad, reaching down to level of sterna. Pore formula usually normal: 5, 7, 9, 10, 12, 13, 15–18(19), porosteles if any very short, ozopores located either between both lateral lobulations or on caudal lobulation near lateral margin. Tergal setae absent. Epiproct fully hidden under penultimate segment. Sterna narrow, much less than length of coxa, deeply impressed along main axis of body, without modifications. Legs relatively short, barely extending beyond lateral edges of paraterga, without modifications; prefemora longest. Gonapophyses on male coxae 2 vestigial. Gonopod aperture subcordiform, broad, nearly or about as wide as male prozona 7. Gonopods rather simple; coxae subglobose, moderately large, microgranulate and microsetose laterally, with usual long cannulae and usually a small, but evident apicoventral tubercle; very long and slender telopodites unusually strongly exposed, partly also due to a small gonocoel, both or only their solenophores (sph) directed mesad and strongly crossing each other; each telopodite deeply biramous, consisting of a shorter, usually more simple solenomere (sl) held either subparallel or subrectangular (directed caudad) to a longer, normally more elaborate sph (Figs 15 & 17D–F). [details]